Todites emarginatus Naugolnykh
Plant Fossil Names Registry Number: PFN001920
Act LSID: urn:lsid:plantfossilnames.org:act:1920
Author: S. V. Naugolnykh
Rank: species
Reference: Naugolnykh, S. V. (2013): Permian Ferns of Western Angaraland. – Paleontological Journal 47(12): 1379–1462., link
Page of description: 1408
Illustrations or figures: pl. 14, figs 1–3, pl. 15, fig. 5
Types
Holotype 4851/173, Geological Institute, Russian Academy of Sciences, Moscow, Russia
Figures: figs 10j, l, 11, 12, pl. 14, figs 1–3, pl. 15, fig. 5
Original diagnosis/description
Fronds at least bipinnate. Sterile pinnules ranging from pecopteroid to sphenopteroid, with lobate margins and round apices. Pinnule midveins slightly undulating. Lateral veins dichotomizing up to two times. Fertile pinnules possessing isolated free sporangia, sometimes arranged in compact clusters. Thick-walled cells specialized for opening stomium located above stomium and forming short row. Each fertile pinnule with 80–100 sporangia.
Etymology
From the Latin emarginatus (emarginate, sinuate), based on the dissected leaf lamina of pinnules into round lobes.
Stratigraphy
Permian
Upper Permian, Severodvinian Stage
Locality
Russian Federation
Pechora Coal Basin, Pechora River, outcrop no. 22 (after Zalessky, 1934a)
Plant fossil remain
macro- and meso-fossils-embryophytes except wood
Comments
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