Actinodaphnoxylon zileensis Akkemik et Mantz. in Akkemik, Mantzouka, Tunç & Koçbulut
Plant Fossil Names Registry Number: PFN001579
Act LSID: urn:lsid:plantfossilnames.org:act:1579
Authors: Ü. Akkemik & D. Mantzouka
Rank: species
Genus: Actinodaphnoxylon Akkemik et Mantz. in Akkemik, Mantzouka, Tunç & Koçbulut
Reference: Akkemik, Ü., Mantzouka, D., Tunç, U. & Koçbulut, F. (2021): The first paleoxylotomical evidence from the Mid-Eocene Climate Optimum from Turkey. – Review of Palaeobotany and Palynology 285: 104356 (20 pp.).
Page of description: 8
Illustrations or figures: fig. 4 (right specimen), pls IV, V, VI
Name is type for
Actinodaphnoxylon Akkemik et Mantz. in Akkemik, Mantzouka, Tunç & Koçbulut 2021
Types
Holotype ZIL02, Department of Forest Botany, Faculty of Forestry, Istanbul University-Cerrahpasa, Istanbul, Turkey
Figures: fig. 4 (right specimen)
Original diagnosis/description
Growth ring boundaries absent, wood diffuse porous, vessels predominantly in radial multiples up to 7 vessels, and clusters present. Ratio of vessels in groups 92%. Solitary vessels very rare (8%). Tyloses common. Pits on vessels alternate, and sometimes scalariform-like pits present in vessel-ray pit areas. Vessel frequency 20-40 per square mm, helical thickenings present, perforation plates simple, axial parenchyma rare, scanty paratracheal and vasicentric, rays 1-3 seriate, oil and/or mucilage cells can be seen mainly at the procumbent and marginal ray cells and among the fibers. Rays heterocellular with one (-two) rows of upright &/or square marginal cells (usually oil and mucilage cells). Dried oil cells look like crystals. Septate fibers present
Etymology
The epithet “zileensis” originates from the name of the city “Zile”, which have the fossil site.
Stratigraphy
Paleogene, Eocene
Çekerek Formation, middle Eocene
Locality
Turkey
Zile-Tokat
Plant fossil remain
fossil wood
Notes
The volcanism ceased in the middle Maastrichtian, and the interval between late Campanian and middle Eocene in Central Pontides is represented by a thick sequence of siliciclastic and calciclastic turbidities. Coeval sequences in the south are shallow marine and are separated by unconformities. The marine deposition in the Central Pontides ended in the Middle Eocene as a consequence of collision of the Pontides with the Kırsehir Massif. The deposition starts when the distal parts of the passive continental margin enter the subduction zone, which leads to the uplift of the active margin. This process leads to the development of retro forearc basins filled with turbidities. A critical stage in the collision is reached when the whole of orogen is lifted above sea level: this corresponds to the end of the Lutetian (Middle Eocene) in the Central Pontides (Okay et al. 2018).
The Eocene volcano-sedimentary successions around the study area are represented by shallow marine sedimentary units at the lower parts and lava flows and volcanoclastic units at the middle to upper parts. The field data from the region provided by Göçmengil et al. (2018) demonstrated that middle Eocene magmatic units along the post-collision zone concomitantly developed in a wide area and triggered the magmatism controlled by the region-scale delamination and/or lithospheric removal processes.
The transgressive sequence consisting of conglomerate, sandstone, sandy limestone, limestone, claystone and marl intercalation, located in the south and north of the Çorum river valley in the study area and seated on the older units with an angular unconformity, was first defined and called as Çekerek Formation by Özcan et al. (1980). The unit crops out widely on the northern and southern slopes of the Çorum stream valley in the study area. Çekerek Formation consists of a transgressive sequence consisting of gray-yellow colored sandstone, claystone, clayey limestone, sandy limestone, limestone, marl alternation, starting with pebbles at the bottom.
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